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Smith, WC; Price, DA; Greenberg, RM; Battelle, BA (1993). "Opsins from the lateral eyes and ocelli of the horseshoe crab, Limulus polyphemus". Proceedings of the National Academy of Sciences of the United States of America. 90 (13): 6150–4. doi:10.1073/pnas.90.13.6150. PMC 46885 Freely accessible. PMID 8327495. 

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